Classification of Santalaceae
Hieronymus (1894, in Engler & Prantl)

Tribe Anthoboleae
1. Anthobolus
2. Exocarpos
3. Champereia (now in Opiliaceae)

Tribe Osyrideae
4. Acanthosyris
5. Buckleya
6. Cervantesia
7. Choretrum
8. Colpoon
9. Comandra
10. Fusanus (now considered part of Santalum)
11. Henslowia (now = Dendrotrophe)
12. Jodina
13. Leptomeria
14. Omphacomeria
15. Osyris
16. Phacellaria
17. Pyrularia
18. Santalum (including Mida)
19. Scleropyrum

Tribe Thesieae
20. Arjona
21. Osyridocarpus
22. Quinchamalium
23. Thesidium
24. Thesium

Not known at time or not recognized as distinct:
Amphorogyne
Daenikera
Dendromyza
Dufrenoya
Geocaulon
Kunkeliella
Mida
Myoschilos
Nanodea
Nestronia
Okoubaka
Rhoiacarpos
Spirogardnera

Engler & Krause (1935)
Loranthaceae ­ Viscoideae ­ Eremolepideae
Antidaphne
Eremolepis
Eubrachion
Lepidoceras

Classification of Santalaceae
Pilger (1935, in Engler and Harms)

Tribe Anthoboleae
1. Anthobolus
2. Exocarpos

Tribe Osyrideae
3. Acanthosyris
4. Buckleya
5. Cervantesia
6. Choretrum
7. Colpoon
8. Comandra
9. Darbya (now =Nestronia)
10. Eucarya (now in Santalum)
11. Geocaulon
12. Henslowia (now = Dendrotrophe)
13. Jodina
14. Leptomeria
15. Mida
16. Myoschilos
17. Nanodea
18. Omphacomeria
19. Osyris
20. Phacellaria
21. Pyrularia
22. Rhoiacarpos
23. Santalum
24. Scleropyrum

Tribe Thesieae
25. Arjona
26. Osyridocarpus
27. Quinchamalium
28. Thesidium
29. Thesium
30. Calyptosepalum, now considered Euphorbiaceae, near Drypetes.

Not known at time or not recognized as distinct:
Amphorogyne
Daenikera
Dendromyza
Dufrenoya (=Hylomyza)
Kunkeliella
Okoubaka
Spirogardnera

Engler and Krause (1935)
Loranthaceae ­ Viscoideae ­ Eremolepideae (in 1982 Eremolepidaceae)
Antidaphne (as well as Eremolepis)
Eubrachion
Lepidoceras

Classification of Santalaceae
Stauffer (1969)


This classification was not actually published as such, but can be assumed from his works.
Genera in bold transferred from Pilger's Osyrideae.

Tribe Anthoboleae
1. Anthobolus
2. Exocarpos
3. Omphacomeria

Tribe Osyrideae
4. Acanthosyris
5. Buckleya
6. Cervantesia
7. Colpoon
8. Comandra
9. Darbya (now =Nestronia)
10. Eucarya (now in Santalum)
11.Geocaulon
12. Jodina
13. Mida
14. Myoschilos
15. Nanodea
16. Okoubaka (transferred from Olacaceae)
17. Osyris
18. Pyrularia
19. Rhoiacarpos
20. Santalum
21. Scleropyrum

Tribe Thesieae
22. Arjona
23. Osyridocarpus
24. Quinchamalium
25. Thesidium
26. Thesium
27. Kunkeliella ­ added by Stearn (1972).

Tribe Amphorogyneae
28. Amphorogyne
29. Choretrum
30. Cladomyza
31. Daenikera
32. Dendromyza
33. Dendrotrophe (previously Henslowia)
34. Dufrenoya (previously Hylomyza)
35. Leptomeria
36. Phacellaria

Loranthaceae ­ Viscoideae ­ Eremolepideae (in 1982 Eremolepidaceae)
Antidaphne
Eubrachion
Lepidoceras

Classification of Santalaceae
Nickrent January 2005

Pyrularia Clade
1. Acanthosyris
2. Cervantesia
3. Jodina
4. Okoubaka
5. Pyrularia
6. Scleropyrum

Santalum Clade
7. Antidaphne
8. Colpoon
9. Eubrachion
10. Exocarpos
11. Lepidoceras
12. Omphacomeria
13. Myoschilos
14. Nestronia
15. Osyris
16. Rhoiacarpos
17. Santalum

Thesium Clade
18. Buckleya
19. Kunkeliella
20. Osyridocarpus
21. Thesium
22. Thesidium

Comandra Clade
23. Comandra
24. Geocaulon

Quinchamalium Clade
25. Arjona
26. Mida
27. Nanodea
28. Quinchamalium

Amphorogyne Clade
29. Amphorogyne
30. Choretrum
31. Daenikera
32. Dendromyza
33. Dendrotrophe
34. Dufrenoya
35. Leptomeria
36. Phacellaria
37. Spirogardnera

Excluded from Santalaceae and placed in Opiliaceae
1. Anthobolus

SIUC / College of Science / Parasitic Plant Connection / Santalaceae
URL: http://www.parasiticplants.siu.edu/Santalaceae/SantClassif.html
Last updated: 04-Jan-05 / dln